Maybe you've got a bad part then.
If you swap the 328s between the 2 boards, does the R2 work and R3 not work?
That was next on my list I'm looking for the tube of 328 I have to swap one and see.
My uno rev3 has had the chip pulled so much i've about broke the pins off lol.
Well I couldn't find the tube of 328s I have so I pulled chip and swaped them they both work now maybe the chip wasn't seated good or something on the Rev2 I guess