Sending the arduino pin HIGH connected to "motor 1 forward" should turn on the motor but it doesn't it turns it off.
Sending that same pin LOW turns the motor on.
According to the datasheet, it should be operating the way you expected it to originally.
The only way it wouldn't is if you had both pins set high to begin with and pulled one line low. But it would move in the opposite direction than you expected in that case.